[Sunred]

So I think it is safe to tell that other motif behind Zeronos as of this episode. Zeronos's other motif aside from Ushiwakamaru and Benkei is the Asian folklore behind the Summer Triangle made up of Altair, Vega, and Deneb, and most importantly the myth behind the Jujly 7th festival celebrated within Asia, known as Tanabata in Japan.
The story is called The Cowherd and the Weaver Girl, Hikoboshi and Orihime in Japan. Some details vary depending on country but the common gist of things is that Orihime was a great weaver and her father who happened to be the ruler of the galaxy found it saddening she was a workaholic. So she introduced her to Hikoboshi the cowherd and both fell in love. However, their love for each other grew so much that they started slacking off and the emperor was angered at how they changed so he banished them to the farthest depths of the galaxy from each other. Orihime was deeply saddened and her tears became the summer rainy season and feeling sorry for her he decided to let her meet Hikoboshi once every year. Altair is Hikoboshi, Vega is Orihime, and the flock of crows and magpies that bands together to form the bridge for them to meet every year is Deneb hence why Deneb Imagin is a crow motif because of this lore and the karasu tengu stuff thrown in the mix.